Coloborhynchus



The pterosaurs dentition rests in the Upper Cretaceous in North Africa are relatively common among the fauna which got preserved from those ecosystems around 100 million years ago. However, only two different pterosaur taxons have been described. Mader & Kellner (1999) (Full reference: B. J. Mader and A. W. A. Kellner. 1999. A new anhanguerid pterosaur from the Cretaceous of Morocco. Boletim do Museu Nacional - Geologia 45:1-11) described the Coloborhynchus moroccensis species, belonging to the Anhangueridae family, which lived at the middle of the Cretaceous (between the Albian and Cenomanian stages, about 105 million years ago). Remains have been discovered in Morocco. Another alternative combination in the nomenclature of this taxon is Siroccopteryx moroccensis.

Phylogeny: The descriptors of Siroccopteryx placed this genus in the family Anhangueridae, sensu Kellner. David Unwin, however, indicated in 2001 that it was a species of Coloborhynchus, then he called it as C. moroccensis and being a member of the Ornithocheiridae. This has been controversial. In the same year, Michael Festnacht suggested it was more similar to Anhanguera due to the wide end of the snout. In 2009, Kellner considered that SiroccopteryxColoborhynchus clavirostris and Uktenadactylus probably formed together a clade within Anhangueridae. [This last paragraph is from Wikipedia - License: Attribution-ShareAlike 3.0 Unported]